South Africa edged England 31-28 in a nail-biting one-off Test match at Twickenham on Saturday.
The Springboks led 13-6 at half-time, Owen Farrell's two penalties cancelled out by an intercept try by centre Jan Serfontein, Pat Lambie kicking the conversion and two penalties.
Cobus Reinach added a second South African try shortly after the break, but England, with Victor Matfield in the sinbin, hit back with a brace of tries from prop David Wilson and replacement No 8 Ben Morgan, both converted by Farrell.
Schalk Burger handed South Africa back the lead, England hooker Dylan Hartley sin-binned for stamping.
Lambie and George Ford traded penalties before the Springbok fly-half hit a 77nd minute drop-goal, Brad Barritt grabbing a late consolation try for the home side.
Elsewhere, Wales beat Fiji 17-13 – also a one-off Test – at Cardiff's Millennium Stadium.
In a disappointinly error-ridden game, the home side scored three first-half tries through wingers George North and Alex Cuthbert, and a penalty try, the latter converted by Rhys Priestland.
Fiji, who saw prop Campese Ma'afu sent off in the 52nd minute after a second yellow card, replied with two Nemani Nadolo penalties, the giant centre grabbing a 78th intercept try he also converted.
