Wales boss Warren Gatland has told his players they are under pressure to perform against World Cup warm-up opponents England on Saturday.
The fierce rivals meet in Cardiff less than a week after Gatland’s men were beaten 33-19 at Twickenham.
Gatland has made just three changes to the line-up in south-west London, and it features 10 players who were on starting duty in a Six Nations title and Grand Slam-clinching victory over Ireland five months ago.
“It’s easy enough to make a lot of changes, but some of the players would have been disappointed with last weekend, and particularly that first 20 minutes,” Gatland said.
“For us, it’s about a chance to redeem ourselves and give those players another hit-out and go out there and perform, and put some pressure on them to perform as well.
“There is a lot of pressure on those guys on the weekend, because if some of them don’t front and don’t perform, they are opening the door for someone else.
“If you win last week and you win this weekend, you pretty much would have sealed, potentially, a starting spot for the first game or first couple of games in the World Cup.
“That’s why we wanted to make sure that players are aware and that they are under pressure.”
Scarlets duo Jonathan and James Davies will become the first set of brothers since 2006 to start for Wales when they feature against England.
Gatland has named openside flanker James, who wins his fourth cap, alongside back-row colleagues Aaron Wainwright and Ross Moriarty, while older sibling Jonathan partners Hadleigh Parkes in midfield.
They are the first set of brothers to play alongside each other for Wales since Jamie and Nicky Robinson 13 years ago.
James Davies replaces Justin Tipuric, while fly-half Dan Biggar takes over from an injured Gareth Anscombe and lock Jake Ball is in for Adam Beard.
“It’s been tough for James,” Gatland added. “He has been really struggling with back issues, and he has dug in in terms of the way he has pushed himself when he has been in a lot of pain.
“It’s a fantastic opportunity for him. He is pretty fired up.
“He is kind of someone who for his whole career people have knocked him and said he’s too small and not able to compete at the highest level.
“We probably had some of the same thoughts as well, but having coached him and been involved with him, we have been incredibly impressed with him.
“We’ve always been looking at him and seeing his potential. He has been unfortunate that for a few campaigns he has had injuries. We were very impressed with him in Argentina last year.”
Biggar starts with Anscombe having suffered a World Cup-ending knee ligament injury at Twickenham.
On Anscombe, Gatland said: “He’s done his ACL (anterior cruciate ligament) and a meniscus as well.
“He did it when he made the break down the sideline. It wasn’t from a tackle, he’s just stepped off his right leg. You can see the jarring on the video.
“He said to the physios he felt he had jarred his knee and stayed on thinking he could run it off.
“He asked the physios if he’d done any more damage, and they said: ‘No, you’d already done that and it wasn’t going to get any worse’.
“Unfortunately, he’s going to require surgery and it’s going to keep him out for most of the season.”
Tipuric (ankle) and Beard (wrist), meanwhile, have been left out as a precaution this weekend.
England have welcomed a string of injuries to their back row as an opportunity to contingency plan in case disaster strikes at the World Cup.
Eddie Jones has taken the unusual step of naming two hookers on the bench for Saturday’s Test against Wales in Cardiff with Jamie George and Jack Singleton supporting Luke Cowan-Dickie.
Singleton won his first cap against the Six Nations Grand Slam champions last weekend and is poised to claim the second as a back row replacement in the World Cup warm-up Test at the Principality Stadium.
Tom Curry, Mark Wilson and Sam Underhill are unavailable through injury, forcing England to examine otherwise unlikely options for the back of the scrum.
“This is actually great for us – it’s the type of thing that could happen to us at the World Cup. Players have to adapt,” forwards coach Steve Borthwick said.
“We’ve got three very good hookers and Jack is there to operate as a utility forward. It’s great opportunity for him and for us to learn to adapt.”
Curry (shoulder), Wilson (ribs) and Underhill (toe) are involved in training and should play some part in the remaining two warm-up games against Ireland and Italy.
Henry Slade, however, might miss all four Tests because of his knee injury and has tentatively been pencilled in to face Italy on September 6 – the same date it is hoped his Exeter colleague Jack Nowell will return from an ankle problem.
Ruaridh McConnochie will make his England debut against Wales a week later than hoped for after overcoming a hip problem.
The Bath wing and sevens specialist was picked in the team to face Warren Gatland’s men at Twickenham last weekend but was among a trio of late withdrawals due to injury.
Attack coach Scott Wisemantel is backing the Rio 2016 silver medallist to thrive despite the cauldron-like atmosphere awaiting in the Welsh capital.
“It’s a great opportunity for him. He deserves his chance and it was unfortunate last week when he had the minor injury blow,” Wisemantel said.
“Cardiff on Saturday will be intense but once he gets going and once the whistle is blown, he’ll just settle into things and will be fine.
“He won’t have any problems. The guy has played on the big stage – the Olympic Games. So it won’t be a problem at all.
“He’ll draw on that experience, but he’ll also be focused on the game. He’ll do the job and do it well.”
Anthony Watson has been left out of the matchday 23 after completing his comeback from a 17-month absence due to an Achilles issues, so Joe Cokanasiga lines up on the opposite wing.
England captain Owen Farrell makes his first appearance of the World Cup warm-up schedule but is confined to a place on the bench, meaning George Ford will lead the team once more.
In total there are three changes to the side that routed Wales 33-19 with Maro Itoje restored to the second row in place of Charlie Ewels with Joe Launchbury still present.
The final adjustment comes in the back row where Courtney Lawes packs down at blindside flanker, forcing Lewis Ludlam to switch to seven.
Piers Francis and Jonathan Joseph are reunited as the centre pairing and New Zealand-born Willi Heinz wins his second cap at scrum-half.
“We want to back up Sunday. We feel there are areas of growth that we can show on Saturday,” Ford said.
“Going to Cardiff is brilliant for us. It’s brilliant for us as players to be challenged like that. We’re coming up against strong opponents so we’ll make the most of it.”
Eddie Jones’ England took the bragging rights at Twickenham last weekend, with their convincing 33-19 victory ending both Wales’ 14-match unbeaten run and their hopes of replacing New Zealand as rugby union’s number one-ranked nation.
Here, PA reviews the last four Cardiff encounters between Wales and England.
Wales 21-13 England, 23 February 2019
England’s Six Nations title and Grand Slam hopes were abruptly ended at the Principality Stadium despite Jones’ side leading at the break.
Tom Curry’s first England try – the visitors’ only one of the fixture – helped guide his side into a 10-3 lead but a determined Wales came roaring back and powered to victory following late tries by lock Cory Hill and wing Josh Adams.
They went on to win the Grand Slam themselves after wins over Scotland and Ireland.
Wales 16-21 England, 11 February 2017
Two years earlier, it was England’s turn to come from behind, trailing by five points at the interval before they snatched an unlikely Six Nations victory.
Liam Williams cut through the England ranks to cancel out Ben Youngs’ earlier try and edge the hosts back in front but thanks to Owen Farrell’s impressive goalkicking, England still had a fighting chance and remained within touching distance.
A critical error then gifted the visitors their golden opportunity to break Welsh hearts with less than five minutes remaining, as centre Jonathan Davies’ poor clearance saw Elliot Daly freed in space and he touched down in the corner.
Wales 16-21 England, 6 February 2015
It was the same scoreline in 2015, although this time it was a far less experienced and injury-ridden England that took the Six Nations spoils.
Wales opened up a substantial early lead as Leigh Halfpenny’s second-minute penalty and Rhys Webb’s try only six minutes later left England shell-shocked, but Anthony Watson helped the visitors hit back by hauling Mike Brown’s grubber kick over the line to reduce the margin.
Jonathan Joseph then bulldozed through two defenders in response to Dan Biggar’s drop-goal, and the comeback was completed when George Ford kicked a penalty before booting another from long range with time running out.
Wales 30-3 England, 16 March 2013
One of Wales’ greatest triumphs over their neighbours, a winner-takes-all Six Nations contest will live long in the memory as Wales’ biggest margin of victory over England.
Three Halfpenny penalties – to Farrell’s single strike – took Wales into a 9-3 lead at the break before the hosts ran away with things as a pair of second-half tries from wing Alex Cuthbert left England’s Grand Slam dream in tatters.
Biggar’s drop goal and penalty put the icing on the cake for Wales, beating their previous best win against England of 25-0 and landing the title in spectacular fashion.